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He earned a doctorate in physiology from the University of London in July 1902, in 1910 he became a Fellow of Trinity College, and an Honorary Fellow egectos Emmanuel College. In 1914 he was elected to the Chair of Biochemistry at Cambridge University and efectps Cambridge students included neurochemistry pioneer Judah Hirsch Quastel and pioneer embryologist Joseph Needham. Hopkins had for a time studied how cells obtain energy via a complex metabolic process of oxidation and reduction reactions. His study in 1907 with Sir Walter Morley Fletcher of the connection between lactic acid and muscle contraction que es viagra professional 100 one of secumdarios achievements of his work on the biochemistry of the cell. He and Fletcher showed that oxygen depletion causes an accumulation of acid in the muscle. Their work paved the way for the discovery by Archibald Hill. This led him to suggest the existence in normal diets of tiny quantities of as yet unidentified substances that are essential for animal growth and these hypothetical substances he called accessory food factors, later efectos secundarios de usar sildenafil viagra vitamins. It was this work that led his being awarded the 1929 Nobel Prize in Physiology silldenafil Medicine, during World War I, Hopkins continued his work on the nutritional value of vitamins. His efforts were especially valuable in a time of food shortages and he agreed to study the nutritional value of margarine and swcundarios that it was, as suspected, inferior to butter because it lacked the vitamins A and D.

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